Institution Program GRE Not Required Program Link
Albert Einstein College of Medicine Biomedical Sciences X https://www.einstein.yu.edu/education/phd/prospective-students/admissions.aspx
Baylor College of Medicine Graduate School of Biomedical Sciences X https://www.bcm.edu/education/schools/graduate-school-of-biomedical-sciences
Boston University Molecular Biology, Cell Biology, & Biochemistry X http://www.bu.edu/mcbb/admissions/
Brandeis University Molecular and Cell Biology X http://www.bio.brandeis.edu/grad/mcb/mcb_phd_rev.html
Brandeis University Neuroscience X http://www.bio.brandeis.edu/grad/neuro/neuro_phd.html
Brown University Ecology and Evolutionary Biology X https://www.brown.edu/graduateprograms/biomed-ecology-and-evolutionary-biology-phd
California State University Dominguez Hills Biology X https://www.csudh.edu/gsr/graduate-studies/graduate-admission/#
California State University Sacramento Master’s of Stem Cel Biology X https://www.csus.edu/bios/graduate%20program/ma-stemcell.html
Carnegie Mellon University Biological Sciences X https://www.cmu.edu/bio/graduate/phd_program/admissions.html
Case Western Reserve University Biomedical Sciences Training Program X https://case.edu/med/BSTP/apply.html
City University of New York Biology X https://www.gc.cuny.edu/Prospective-Current-Students/Prospective-Students/Admissions#!
Cold Spring Harbor Laboratory PhD Program X https://www.cshl.edu/education/phd-program/how-to-apply/
Cornell University Biochemistry, Molecular, and Cell Biology X https://bmcb.cornell.edu/
Cornell University Biomedical Engineering X https://www.bme.cornell.edu/bme/programs/graduate-programs
Cornell University Genetics, Genomics, and Development X https://gendev.cornell.edu/
Dartmouth Molecular & Cell Biology X https://graduate.dartmouth.edu/mcb/
Dartmouth Program in Experimental & Molecular Medicine X https://graduate.dartmouth.edu/pemm/
DePaul University Biological Sciences (MS) X website
Duke University Biochemistry X https://gradschool.duke.edu/admissions/apply/gre-scores
Duke University Cell & Molecular Biology X https://gradschool.duke.edu/admissions/apply/gre-scores
Duke University Developmental & Stem Cell Biology X https://gradschool.duke.edu/admissions/apply/gre-scores
Duke University Immunology X https://gradschool.duke.edu/admissions/apply/gre-scores
Duke University Molecular Cancer Biology X https://gradschool.duke.edu/admissions/apply/gre-scores
Duke University Molecular Genetics and Microbiology X https://gradschool.duke.edu/admissions/apply/gre-scores
Duke University Pharmacology X https://gradschool.duke.edu/admissions/apply/gre-scores
Eastern Virginia Medical School Biomedical Sciences X https://www.evms.edu/education/doctoral_programs/biomedical_sciences/
Emory University Biological and Biomedical Sciences (except Population Bio) X http://www.biomed.emory.edu/
Emory University Population Biology, Ecology and Evolution X http://www.biomed.emory.edu/PROGRAM_SITES/PBEE/index.html
Georgetown School of Medicine Pharmacology and Physiology X https://pharmacology.georgetown.edu/phd_applying
Gerstner Sloan Kettering Biomedical Sciences X https://www.sloankettering.edu/gerstner/admissions/apply
Harvard University Biological and Biomedical Sciences X http://www.hms.harvard.edu/dms/bbs/
Harvard University Biological Sciences in Dental Medicine X https://gsas.harvard.edu/programs-of-study/all/biological-sciences-dental-medicine
Harvard University Biological Sciences in Public Health X https://www.hsph.harvard.edu/biological-sciences/
Harvard University Biological Sciences in Public Health X https://www.hsph.harvard.edu/biological-sciences/
Harvard University Biophysics X https://gsas.harvard.edu/programs-of-study/all/biophysics
Harvard University Chemical Biology X http://chembiophd.harvard.edu/
Harvard University Immunology X http://dms.hms.harvard.edu/immunology/index.html
Harvard University Molecular and Cellular Biology X https://gsas.harvard.edu/programs-of-study/all/molecular-and-cellular-biology
Harvard University Neuroscience X http://dms.hms.harvard.edu/neuroscience/index.html
Harvard University Organismic & Evolutionary Biology X https://gsas.harvard.edu/programs-of-study/all/organismic-and-evolutionary-biology
Harvard University Systems Biology X http://sysbiophd.harvard.edu/prospective-students/admissions
Harvard University Virology X http://dms.hms.harvard.edu/virology/index.html
Icahn School of Medicine at Mount Sinai Biomedical Science, Neuroscience X http://icahn.mssm.edu/education/phd
Indiana University Biology X https://www.bio.indiana.edu/
Indiana University Cell, Molecular and Cancer Biology X website
Indiana University School of Medicine Biomedical X https://medicine.iu.edu/education/graduate-degrees/phd/admissions/
Iowa State University Ecology and Evolutionary Biology X https://www.eeb.iastate.edu/admissions
Johns Hopkins School of Medicine Biochemistry, Cellular & Molecular Biology X http://bcmb.bs.jhmi.edu/
Johns Hopkins School of Medicine Cellular and Molecular Medicine Program X http://cmm.jhmi.edu/index.php/application-information/
Johns Hopkins School of Medicine Molecular Biophysics X http://pmb.jhu.edu/
Johns Hopkins School of Medicine Neuroscience X http://neuroscience.jhu.edu/graduate/apply
Massachusetts Institute of Technology Biology X https://biology.mit.edu/graduate/prospective-students/application-process/
Medical University of South Carolina Biomedical Sciences X www.musc.edu/grad
Michigan Tech Biological Sciences X http://www.mtu.edu/gradschool/programs/degrees/biological-sciences/
New York University Biology X https://as.nyu.edu/biology/programs/phd.html
New York University Sackler Institute Biomedical Sciences X https://med.nyu.edu/research/sackler-institute-graduate-biomedical-sciences/
NIH Oxford-Cambridge Scholars Program X https://oxcam.gpp.nih.gov/
Northwestern University Life Sciences X Website
Ohio State University Biochemistry X https://osbp.osu.edu/admissions
Ohio State University Evolution, Ecology and Organismal Biology X https://eeob.osu.edu/grad/graduate-program
Ohio State University Microbiology X https://microbiology.osu.edu/gradstudents/admissions
Penn State College of Medicine Anatomy X https://med.psu.edu/anatomy-phd
Penn State College of Medicine Biomedical Sciences X https://med.psu.edu/biomedical-sciences-phd
Penn State College of Medicine Neuroscience X https://med.psu.edu/neuroscience-phd
Penn State University Park Biochemistry, Microbiology, and Molecular Biology X http://bmb.psu.edu/graduate
Princeton University Molecular Biology X https://molbio.princeton.edu/
Princeton University Neuroscience X http://pni.princeton.edu/
Purdue University Chemistry X https://www.chem.purdue.edu/faq/index.php#gre
Rockefeller University Graduate Program X https://graduateapplication.rockefeller.edu/
Rutgers University Biochemistry (Molecular Biosciences) X https://molbiosci.rutgers.edu/
Rutgers University Cell and Developmental Biology (Molecular Biosciences) X https://molbiosci.rutgers.edu/
Rutgers University Microbiology & Molecular Genetics (Molecular Biosciences) X https://molbiosci.rutgers.edu/
Rutgers University Neuroscience X http://rwjms.rutgers.edu/gsbs/programs/neuroscience/index.html
Rutgers University Pharmacology, Cellular & Molecular (Molecular Biosciences) X https://molbiosci.rutgers.edu/
Rutgers University Physiology & Integrative Biology (Molecular Biosciences) X https://molbiosci.rutgers.edu/
Rutgers University (Newark) Cell Biology, Neuroscience and Physiology (Newark) X website
Rutgers University (Newark) Immunity and Inflammation (Newark) X https://njms-web.njms.rutgers.edu/gsbs/multidisciplinary/multidisciplinary_i3.php
Rutgers University (Newark) Molecular Biology Genetics and Cancer (Newark) X https://njms-web.njms.rutgers.edu/gsbs/multidisciplinary/multidisciplinary_cbnp2.php
San Francisco State University Biology (Master’s) X http://biology.sfsu.edu/content/applying-sfsu-biology-graduate-programs
Stanford University Biosciences Varies by program https://biosciences.stanford.edu/prospective-students/how-to-apply/
Stowers Institute for Medical Research Graduate Program X http://www.stowers.org/gradschool/admissions
Texas A&M University Ecology and Evolutionary Biology X https://eeb.tamu.edu/
Thomas Jefferson University Graduate School of Biomedical Sciences X https://www.jefferson.edu/university/life-sciences.html
University of Alabama Birmingham Biomedical Sciences X http://www.uab.edu/gbs/home/admissions
University of Arizona Biological and Biomedical Sciences X https://abbs.arizona.edu/
University of Buffalo Program in Biomedical Science (PPBS) X http://medicine.buffalo.edu/phdprogram.html
University of Buffalo Microbiology and Immunology X http://medicine.buffalo.edu/departments/micro/education/phd.html
University of California Berkeley Environmental Science, Policy, and Management X https://ourenvironment.berkeley.edu/graduate-programs/faq
University of California Berkeley Integrative Biology (IB) X http://ib.berkeley.edu/
University of California Berkeley Microbiology X http://plantandmicrobiology.berkeley.edu/graduate-program-microbiology
University of California Berkeley Molecular and Cell Biology X https://mcb.berkeley.edu/grad/prospective-graduates/admissions-and-recruitment
University of California Berkeley Neuroscience X https://neuroscience.berkeley.edu/ph-d-program/
University of California Berkeley Plant Biology X http://plantandmicrobiology.berkeley.edu/plant-biology-0
University of California Davis Biochemistry, Molecular, Cellular and Developmental Biology X http://bmcdb.ucdavis.edu/admissions/index.html
University of California Davis Medical Microbiology & Immunology X https://health.ucdavis.edu/medmicro/
University of California Davis Molecular, Cellular, and Integrative Physiology X https://mcip.ucdavis.edu/
University of California Los Angeles Bioscience (Varies by program) X http://bioscience.ucla.edu/application-procedure
University of California Los Angeles Ecology and Evoltionary Biology X https://www.eeb.ucla.edu/grad_phd.php
University of California San Diego Biomedical Sciences X http://biomedsci.ucsd.edu/how-to-apply/index.html
University of California San Francisco Biomedical Sciences X https://bms.ucsf.edu/
University of California San Francisco Chemistry and Chemical Biology X https://ccb.ucsf.edu/admissions
University of California San Francisco Developmental and Stem Cell Biology X https://dscb.ucsf.edu/
University of California San Francisco Integrative Program in Quantitative Biology X https://ipqb.ucsf.edu/
University of California San Francisco Neuroscience X http://neurograd.ucsf.edu/about-neuroscience-graduate-program
University of California San Francisco Oral and Craniofacial Sciences X https://dentistry.ucsf.edu/programs/oral-cranio-phd
University of California San Francisco Pharmaceutical Sciences and Pharmacogenomics X https://pspg.ucsf.edu/
University of California San Francisco Rehabilitation Science X https://ptrehab.ucsf.edu/rehabilitation-science-phd
University of California San Francisco Tetrad X https://tetrad.ucsf.edu/admissions
University of California Santa Cruz Molecular, Cell, and Developmental Biology X https://pbse.ucsc.edu/mcd/index.html
University of Chicago Bioisciences Programs X https://biosciences.uchicago.edu/programs
University of Chicago Ecology and Evolution X https://pondside.uchicago.edu/ee/index.html
University of Colorado Anschutz Medical Campus Biomedical Sciences X https://medschool.cuanschutz.edu/departments-centers-institutes
University of Colorado Anschutz Medical Campus Cancer Biology X https://medschool.cuanschutz.edu/departments-centers-institutes
University of Colorado Anschutz Medical Campus Cell Biology, Stem Cells and Development X https://medschool.cuanschutz.edu/departments-centers-institutes
University of Colorado Anschutz Medical Campus Immunology X https://medschool.cuanschutz.edu/departments-centers-institutes
University of Colorado Anschutz Medical Campus Microbiology X https://medschool.cuanschutz.edu/departments-centers-institutes
University of Colorado Anschutz Medical Campus Molecular Biology X https://medschool.cuanschutz.edu/departments-centers-institutes
University of Colorado Anschutz Medical Campus Pharaceutical Sciences X https://medschool.cuanschutz.edu/departments-centers-institutes
University of Colorado Anschutz Medical Campus Pharmacology X https://medschool.cuanschutz.edu/departments-centers-institutes
University of Colorado Anschutz Medical Campus Toxicology X https://medschool.cuanschutz.edu/departments-centers-institutes
University of Colorado at Boulder Ecology and Evolutionary Biology X https://www.colorado.edu/ebio/
University of Connecticut Molecular and Cell Biology X https://mcb.uconn.edu/graduate-application-aid/
University of Georgia Integrated Life Sciences X https://ils.uga.edu/
University of Illinois at Urbana-Champaign Molecular and Cellular Biology X https://mcb.illinois.edu/graduate/gradprospect/
University of Iowa Neuroscience X https://neuroscience.grad.uiowa.edu/
University of Kansas Ecology and Evolutionary Biology X https://eeb.ku.edu/how-apply
University of Kansas Molecular Biosciences X https://molecularbiosciences.ku.edu/
University of Kansas Medical Center Interdisciplinary Graduate Program in Biomedical Sciences X http://www.kumc.edu/igpbs.html
University of Maryland Baltimore Varies by program (see link) X https://lifesciences.umaryland.edu/prospective/Admissions/
University of Massachusetts Amherst Molecular and Cellular Biology X http://gpls.cns.umass.edu/mcb
University of Massachusetts Amherst Neuroscience and Behavior X http://gpls.cns.umass.edu/nsb
University of Massachusetts Amherst Plant Biology X http://gpls.cns.umass.edu/pb
University of Massachusetts Medical School Biomedical Sciences X https://www.umassmed.edu/gsbs/
University of Michigan Biophysics X https://lsa.umich.edu/biophysics/graduates/admissions.html
University of Michigan Chemical Biology X http://www.chembio.umich.edu/
University of Michigan Ecology and Evolutionary Biology X https://lsa.umich.edu/eeb
University of Michigan Program in Biomedical Science X website
University of Minnesota Ecology and Evolutionary Biology X https://cbs.umn.edu/academics/departments/eeb/graduate/graduate-admissions
University of Minnesota Biochemistry, Molecular Biology & Biophysics X https://www.med.umn.edu/education-training/graduate-programs
University of Minnesota Integrative Biology and Physiology X https://www.physiology.umn.edu/degrees-and-programs/graduate-program
University of Minnesota Microbiology, Immunology and Cancer Biology X http://www.micab.umn.edu/
University of Minnesota Molecular, Cellular, Developmental Biology & Genetics X https://www.med.umn.edu/education-training/graduate-programs
University of Minnesota Neuroscience X https://med.umn.edu/neuroscience
University of Minnesota Pharmacology X https://www.med.umn.edu/education-training/graduate-programs
University of Missouri Division of Biological Sciences X http://biology.missouri.edu/graduate-studies/apply/
University of Montana Organismal Biology, Ecology, and Evolution X http://hs.umt.edu/dbs/grad-programs/OBE/default.php
University of Nebraska Medical Center Interdisciplinary Graduate Program in Biomedical Sciences X https://www.unmc.edu/gradstudies/programs/phd/interdisciplinary-biomedical-sciences.html
University of New Mexico Biology X http://biology.unm.edu/graduate/graduate-admissions.shtml
University of North Carolina at Chapel Hill Biological and Biomedical Sciences Program X http://bbsp.unc.edu/admissions/
University of Oklahoma Health Sciences Graduate Program in Biomedical Sciences X website
University of Oregon Biology X https://biology.uoregon.edu/graduate-studies/apply/
University of Pennsylvania Biology X https://www.bio.upenn.edu/graduate/
University of Pennsylvania Biomedical Graduate Studies X http://www.med.upenn.edu/bgs/
University of Pittsburgh Biological Sciences X http://www.biology.pitt.edu/graduate/how-apply
University of Pittsburgh Biomedical Informatics X https://www.dbmi.pitt.edu/content/doctoral-degree-0
University of Pittsburgh Cell Biology and Molecular Physiology X website
University of Pittsburgh Cellular and Molecular Pathology X http://www.gradbiomed.pitt.edu/programs/cellular-molecular-pathology
University of Pittsburgh Clinical and Translational Science X http://www.icre.pitt.edu/phd/index.html
University of Pittsburgh Computational Biology X http://www.compbio.cmu.edu/
University of Pittsburgh Integrative Systems Biology X https://www.isb.pitt.edu/
University of Pittsburgh Microbiology and Immunology X http://pmi.pitt.edu/
University of Pittsburgh Molecular Biophysics & Structural Biology X http://www.mbsb.pitt.edu/
University of Pittsburgh Molecular Genetics & Developmental Biology X http://www.gradbiomed.pitt.edu/programs/molecular-genetics-development-biology
University of Pittsburgh Molecular Pharmacology X http://www.gradbiomed.pitt.edu/programs/molecular-pharmacology
University of Pittsburgh Neurobiology X http://cnup.pitt.edu/training/graduate-program/phd-overview
University of South Dakota School of Medicine Basic Biomedical Sciences X https://www.usd.edu/medicine/basic-biomedical-sciences
University of Tennessee Ecology and Evolutionary Biology X https://eeb.utk.edu/
University of Tennessee Microbiology X https://micro.utk.edu/
University of Texas MD Anderson Cancer Center UTHealth Graduate School of Biomedical Sciences X https://gsbs.uth.edu/
University of Texas Southwestern Medical School Graduate School of Biomedical Sciences (Umbrella PhD and MS) X http://www.utsouthwestern.edu/education/graduate-school/application-and-admissions/
University of Vermont Biology X https://www.uvm.edu/cas/biology/application-finances-support
University of Vermont Cellular, Molecular, and Biomedical Sciences X https://www.uvm.edu/cmb
University of Virginia Biomedical Sciences X http://bims.virginia.edu/
University of Virginia Biology X http://bio.as.virginia.edu
University of Virginia Ecology and Evolutionary Biology X http://eeb.virginia.edu
University of Washington Biology X https://www.biology.washington.edu/programs/graduate/admissions
University of Washington Genome Sciences X https://www.gs.washington.edu/academics/gradprogram/index.htm
University of Washington Immunology X http://www.immunology.washington.edu/graduate-program/admissions/admissions-faqs/
University of Washington Molecular Medicine and Mechanisms of Disease X http://www.pathology.washington.edu/academics/mmtp/phd-program/
University of Washington Molecular and Cellular Biology X https://depts.washington.edu/mcb/
University of Wisconsin-Madison Cell and Molecular Biology X https://cmb.wisc.edu/admissions/
University of Wisconsin-Madison Endocrinology X https://www.med.wisc.edu/education/graduate-programs/biomedical-graduate-program-consortium/application-checklist/
University of Wisconsin-Madison Integrated Program in Biochemistry X https://ipib.wisc.edu/
University of Wisconsin-Madison Microbiology X https://microbiology.wisc.edu/ps_apply.php
University of Wisconsin-Madison Neuroscience X https://ntp.neuroscience.wisc.edu/
University of Wisconsin-Madison Nutritional Sciences X https://nutrisci.wisc.edu/graduate/
University of Wisconsin-Madison Pharmacology X https://www.med.wisc.edu/education/graduate-programs/biomedical-graduate-program-consortium/application-checklist/
Vanderbilt University Biological Sciences X https://as.vanderbilt.edu/biosci/graduate/admission.php
Vanderbilt University Interdisciplinary Graduate Program X https://medschool.vanderbilt.edu/igp/
Virginia Commonwealth University Biomedical Sciences Doctoral Portal X https://medschool.vcu.edu/education/bsdp/
Virginia Commonwealth University Clinical and Translational Research, Cancer and Molecular Medicine X
Virginia Tech Biological Sciences X https://www.biol.vt.edu/
Washington State University Biological Systems Engineering X https://bsyse.wsu.edu/graduate-program/how-to-apply/
Washington University in St. Louis DBBS – Biochemistry, Biophysics & Structural Biology X http://dbbs.wustl.edu/divprograms/biophysics/Pages/BBSB.aspx
Washington University in St. Louis DBBS – Developmental, Regenerative & Stem Cell Biology X http://www.dbbs.wustl.edu/divprograms/devbio/Pages/default.aspx
Washington University in St. Louis DBBS – Computational & Systems Biology X
Washington University in St. Louis DBBS – Evolution, Ecology & Population Biology X http://www.dbbs.wustl.edu/divprograms/eepb/Pages/default.aspx
Washington University in St. Louis DBBS – Human & Statistical Genetics X http://dbbs.wustl.edu/divprograms/hsg/Pages/default.aspx
Washington University in St. Louis DBBS – Immunology X http://dbbs.wustl.edu/divprograms/immunology/Pages/default.aspx
Washington University in St. Louis DBBS – Molecular Genetics & Genomics X http://www.dbbs.wustl.edu/divprograms/genetics/Pages/default.aspx
Washington University in St. Louis DBBS – Molecular Microbiology & Microbial Pathogenesis X http://www.dbbs.wustl.edu/divprograms/micro/Pages/default.aspx
Washington University in St. Louis DBBS – Neurosciences X http://dbbs.wustl.edu/divprograms/neuro/Pages/default.aspx
Washington University in St. Louis DBBS – Plant & Microbial Biosciences X http://dbbs.wustl.edu/divprograms/PlantMicroBioSci/Pages/default.aspx
Washington University in St. Louis DDBS – Molecular Cell Biology X http://www.dbbs.wustl.edu/divprograms/cellbio/Pages/default.aspx
Weill Cornell Medicine All biomedical programs except Pharmacology X https://gradschool.weill.cornell.edu/admissions/apply-online
Yale University Combined Program in Biological & Biomedical Sciences X https://medicine.yale.edu/bbs/
Updated August 30, 2019

Please email Joshua Hall (jdhall@unc.edu) or tweet to @jdhallphd for additions or corrections to this list.

Updated August 30, 2019

Reference:

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